<< Chapter < Page Chapter >> Page >

Vascular tissue

The xylem and phloem that make up the vascular tissue of the stem are arranged in distinct strands called vascular bundles, which run up and down the length of the stem. When the stem is viewed in cross section, the vascular bundles of dicot stems are arranged in a ring. In plants with stems that live for more than one year, the individual bundles grow together and produce the characteristic growth rings. In monocot stems, the vascular bundles are randomly scattered throughout the ground tissue ( [link] ).

 Part A is cross section of a dicot stem. In the center of the stem is ground tissue. Symmetrically arranged near the outside of the stem are egg-shaped vascular bundles; the narrow end of the egg points inward. The inner part of the vascular bundle is xylem tissue, and the outer part is sclerenchyma tissue. Sandwiched between the xylem and sclerenchyma is the phloem. Part B is a cross section of a monocot stem. In the monocot stem, the vascular bundles are scattered throughout the ground tissue. The bundles are smaller than in the dicot stem, and distinct layers of xylem, phloem and sclerenchyma cannot be discerned.
In (a) dicot stems, vascular bundles are arranged around the periphery of the ground tissue. The xylem tissue is located toward the interior of the vascular bundle, and phloem is located toward the exterior. Sclerenchyma fibers cap the vascular bundles. In (b) monocot stems, vascular bundles composed of xylem and phloem tissues are scattered throughout the ground tissue.

Xylem tissue has three types of cells: xylem parenchyma, tracheids, and vessel elements. The latter two types conduct water and are dead at maturity. Tracheids are xylem cells with thick secondary cell walls that are lignified. Water moves from one tracheid to another through regions on the side walls known as pits, where secondary walls are absent. Vessel elements are xylem cells with thinner walls; they are shorter than tracheids. Each vessel element is connected to the next by means of a perforation plate at the end walls of the element. Water moves through the perforation plates to travel up the plant.

Phloem tissue is composed of sieve-tube cells, companion cells, phloem parenchyma, and phloem fibers. A series of sieve-tube cells (also called sieve-tube elements) are arranged end to end to make up a long sieve tube, which transports organic substances such as sugars and amino acids. The sugars flow from one sieve-tube cell to the next through perforated sieve plates, which are found at the end junctions between two cells. Although still alive at maturity, the nucleus and other cell components of the sieve-tube cells have disintegrated. Companion cells are found alongside the sieve-tube cells, providing them with metabolic support. The companion cells contain more ribosomes and mitochondria than the sieve-tube cells, which lack some cellular organelles.

Ground tissue

Ground tissue is mostly made up of parenchyma cells, but may also contain collenchyma and sclerenchyma cells that help support the stem. The ground tissue towards the interior of the vascular tissue in a stem or root is known as pith    , while the layer of tissue between the vascular tissue and the epidermis is known as the cortex    .

Growth in stems

Growth in plants occurs as the stems and roots lengthen. Some plants, especially those that are woody, also increase in thickness during their life span. The increase in length of the shoot and the root is referred to as primary growth    , and is the result of cell division in the shoot apical meristem. Secondary growth is characterized by an increase in thickness or girth of the plant, and is caused by cell division in the lateral meristem. [link] shows the areas of primary and secondary growth in a plant. Herbaceous plants mostly undergo primary growth, with hardly any secondary growth or increase in thickness. Secondary growth or “wood” is noticeable in woody plants; it occurs in some dicots, but occurs very rarely in monocots.

Questions & Answers

what is biology
lilian Reply
what is biology
Dada Reply
The scientific study of life.
the virus that causes mumps in humans is composed of a protein outer Shell containing a core of DNA
Daniel Reply
Basic science and applied science question about cancer
Joyce Reply
what are the importance of ATPs
Olatunji Reply
How can biology be studied from a microscopic approach to a global approach
Joyce Reply
The large central opening in the poriferan body is called
Chynna Reply
You go for a long walk on a hot day. Give an example of a way in which homeostasis keeps your body healthy.
Joyce Reply
You sweat.
sweating is your bodies way of keeping you from overheating.
Thank you
what is biology
Neya Reply
biology is the study of life
Biology is the study of Life
is the branch of science which deals with the of living things.
what is metabolism
Sangam Reply
hi palz, what's the ?
Gbolajobi Reply
hi plz, what's the.
pls come a again
Describe the steps and results of reintroducing wolves to Yellowstone National Park.
Natalia Reply
tag and release wolves into Yellowstone. wolves eventually reproduce and the pack grows. as wolves hunt they cull the sick and weak prey. the carcass that is left provides food for other species (scavengers and insect.. etc). this heals the circle of life and contributes to the biodiversity...
before you know it species that are critical to the eco system return. having apex predators is crucial to an ecosystem... it helps run the deer and elk , etc around.
example: there was a species of shrub/ plant that grows along river banks that moose love to eat.. the moose have no predator so they decimate that food source which also helps prevent erosion. when the wolves were reintroduced this changed. oddly enough this plant species started to repopulate in
the areas where wolf feces sat and decayed
which of the following statements about the parts of an egg are false?
Israel Reply
Monotremes include...?
medicinal plants including microbs
vijay Reply
,medicinal plants including microbes
what is biology
Siyanbola Reply
study of living organisms...
study of plants and animals

Get the best Biology course in your pocket!

Source:  OpenStax, Biology. OpenStax CNX. Feb 29, 2016 Download for free at http://cnx.org/content/col11448/1.10
Google Play and the Google Play logo are trademarks of Google Inc.

Notification Switch

Would you like to follow the 'Biology' conversation and receive update notifications?